The 1915 locust plague, which lasted from March to October 1915, was a plague of locusts that stripped areas in and around Palestine of almost all vegetation. This invasion of awesome proportions seriously compromised the already-depleted food supply of the region and sharpened the misery of all Jerusalemites. Midhat Bay, who was the official appointed to fight the plague, helped enact a law which required every male between aged 15 and 60 in cities to collect 20 kilograms of locust eggs or pay a fine of £4.40. Research at Oxford University has identified that swarming behaviour is a response to overcrowding. Increased tactile stimulation of the hind legs causes an increase in levels of serotonin. This causes the locust to change color, eat much more, and breed much more easily. The transformation of the locust to the swarming variety is induced by several contacts per minute over a four-hour period. It is estimated that the largest swarms have covered hundreds of square miles and consisted of many billions of locusts.